Next after or at the back of the mouth comes a long and narrow oesophagus, and close after that a crop or craw, large and spherical, like that of a bird; then comes the stomach, like the fourth stomach in ruminants; and the shape of it resembles the spiral convolution in the trumpet-shell; from the stomach there goes back again, in the direction of the mouth, thin gut, and the gut is thicker than the oesophagus. (See diagram.) Molluscs have no viscera, but they have what is called a mytis, and on it a vessel containing a thick black juice; in the sepia or cuttle-fish this vessel is the largest, and this juice is most abundant. All molluscs, when frightened, discharge such a juice, but the discharge is most copious in the cuttle-fish. The mytis, then, is situated under the mouth, and the oesophagus runs through it; and down below at the point to which the gut extends is the vesicle of the black juice, and the animal has the vesicle and the gut enveloped in one and the same membrane, and by the same membrane, same orifice discharges both the black juice and the residuum. The animals have also certain hair-like or furry growths in their bodies.
In the sepia, the teuthis, and the teuthus the hard parts are within, towards the back of the body; those parts are called in one the sepium, and in the other the ‘sword’. They differ from one another, for the sepium in the cuttle-fish and teuthus is hard and flat, being a substance intermediate between bone and fishbone, with (in part) a crumbling, spongy texture, but in the teuthis the part is thin and somewhat gristly. These parts differ from one another in shape, as do also the bodies of the animals.
The octopus has nothing hard of this kind in its interior, but it has a gristly substance round the head, which, if the animal grows old, becomes hard.
The females differ from the males. The males have a duct in under the oesophagus, extending from the mantle-cavity to the lower portion of the sac, and there is an organ to which it attaches, resembling a breast; 1083
(see diagram) in the female there are two of these organs, situated higher up; (see diagram) with both sexes there are underneath these organs certain red formations. The egg of the octopus is single, uneven on its surface, and of large size; the fluid substance within is all uniform in colour, smooth, and in colour white; the size of the egg is so great as to fill a vessel larger than the creature’s head. The sepia has two sacs, and inside them a number of eggs, like in appearance to white hailstones. For the disposition of these parts I must refer to my anatomical diagrams.
The males of all these animals differ from the females, and the difference between the sexes is most marked in the sepia; for the back of the trunk, which is blacker than the belly, is rougher in the male than in the female, and in the male the back is striped, and the rump is more sharply pointed.
There are several species of the octopus. One keeps close to the surface, and is the largest of them all, and near the shore the size is larger than in deep water; and there are others, small, variegated in colour, which are not articles of food. There are two others, one called the hele-done, which differs from its congeners in the length of its legs and in having one row of suckers-all the rest of the molluscs having two,-the other nicknamed variously the bolitaina or the ‘onion,’ and the ozolis or the ‘stinkard’.
There are two others found in shells resembling those of the testaceans. One of them is nicknamed by some persons the nautilus or the pontilus, or by others the ‘polypus’ egg’; and the shell of this creature is something like a separate valve of a deep scallop-shell. This polypus lives very often near to the shore, and is apt to be thrown up high and dry on the beach; under these circumstances it is found with its shell detached, and dies by and by on dry land. These polypods are small, and are shaped, as regards the form of their bodies, like the bolbidia. There is another polypus that is placed within a shell like a snail; it never comes out of the shell, but lives inside the shell like the snail, and from time to time protrudes its feelers.
So much for molluscs.
<
div id="section58" class="section" title="2"> 2
With regard to the Malacostraca or crustaceans, one species is that of the crawfish, and a second, resembling the first, is that of the lobster; the lobster differing from the crawfish in having claws, and in a few other 1084
respects as well. Another species is that of the carid, and another is that of the crab, and there are many kinds both of carid and of crab.
Of carids there are the so-called cyphae, or ‘hunch-backs’, the crangons, or squillae, and the little kind, or shrimps, and the little kind do not develop into a larger kind.
Of the crab, the varieties are indefinite and incalculable. The largest of all crabs is one nicknamed Maia, a second variety is the pagarus and the crab of Heracleotis, and a third variety is the fresh-water crab; the other varieties are smaller in size and destitute of special designations. In the neighbourhood of Phoenice there are found on the beach certain crabs that are nicknamed the ‘horsemen’, from their running with such speed that it is difficult to overtake them; these crabs, when opened, are usually found empty, and this emptiness may be put down to insufficiency of nutriment. (There is another variety, small like the crab, but resembling in shape the lobster.) All these animals, as has been stated, have their hard and shelly part outside, where the skin is in other animals, and the fleshy part inside; and the belly is more or less provided with lamellae, or little flaps, and the female here deposits her spawn.
The crawfishes have five feet on either side, including the claws at the end; and in like manner the crabs have ten feet in all, including the claws. Of the carids, the hunch-backed, or prawns, have five feet on either side, which are sharp-pointed-those towards the head; and five others on either side in the region of the belly, with their extremities flat; they are devoid of flaps on the under side such as the crawfish has, but on the back they resemble the crawfish. (See diagram.)It is very different with the crangon, or squilla; it has four front legs on either side, then three thin ones close behind on either side, and the rest of the body is for the most part devoid of feet. (See diagram.) Of all these animals the feet bend out obliquely, as is the case with insects; and the claws, where claws are found, turn inwards. The crawfish has a tail, and five fins on it; and the round-backed carid has a tail and four fins; the squilla also has fins at the tail on either side. In the case of both the hump-backed carid and the squilla the middle art of the tail is spinous: only that in the squilla the part is flattened and in the carid it is sharp-pointed. Of all animals of this genus the crab is the only one devoid of a rump; and, while the body of the carid and the crawfish is elongated, that of the crab is rotund.
In the crawfish the male differs from the female: in the female the first foot is bifurcate, in the male it is undivided; the belly-fins in the female are large and overlapping on the neck, while in the male they are smaller 1085
and do not overlap; and, further, on the last feet of the male there are spur-like projections, large and sharp, which projections in the female are small and smooth. Both male and female have two antennae in front of the eyes, large and rough, and other antennae underneath, small and smooth. The eyes of all these creatures are hard and beady, and can move either to the inner or to the outer side. The eyes of most crabs have a similar facility of movement, or rather, in the crab this facility is developed in a higher degree. (See diagram.)
The lobster is all over grey-coloured, with a mottling of black. Its under or hinder feet, up to the big feet or claws, are eight in number; then come the big feet, far larger and flatter at the tips than the same organs in the crawfish; and these big feet or claws are exceptional in their structure, for the right claw has the extreme flat surface long and thin, while the left claw has the corresponding surface thick and round. Each of the two claws, divided at the end like a pair of jaws, has both below and above a set of teeth: only that in the right claw they are all small and saw-shaped, while in the left claw those at the apex are saw-shaped and those within are molar-shaped, these latter being, in the under part of the cleft claw, four teeth close together, and in the upper part three teeth, not close together. Both right and left claws have the upper part mobile, and bring it to bear against the lower one, and both are curved like bandy-legs, being thereby adapted for apprehension and constriction. Above the two large claws come two others, covered with hair, a little underneath the mouth; and underneath these the gill-like formations in the region of the mouth, hairy and numerous. These organs the animal keeps in perpetual motion; and the two hairy feet it bends and draws in towards its mouth. The feet near the mouth are furnished also with delicate outgrowing appendages. Like the crawfish, the lobster has two teeth, or mandibles, and above these teeth are its antennae, long, but shorter and finer by far than those of the crawfish, and then four other antennae similar in shape, but shorter and finer than the others. Over these antennae come the eyes, small and short, not large like the eyes of the crawfish. Over the eyes is a peaky rough projection like a forehead, larger than the same part in the crawfish; in fact, the frontal part is more pointed and the thorax is much broader in the lobster than in the crawfish, and the body in general is smoother and more full of flesh. Of the eight feet, four are bifurcate at the extremities, and four are undivided. The region of the so-called neck is outwardly divided into five divisions, and sixthly comes the flattened portion at the end, and this portion has five flaps, or tail-fins; and the inner or under parts, into which the female 1086
drops her spawn, are four in number and hairy, and on each of the aforesaid parts is a spine turned outwards, short and straight. The body in general and the region of the thorax in particular are smooth, not rough as in the crawfish; but on the large claws the outer portion has larger spines. There is no apparent difference between the male and female, for they both have one claw, whichever it may be, larger than the other, and neither male nor female is ever found with both claws of the same size.
All crustaceans take in water close by the mouth. The crab discharges it, closing up, as it does so, a small portion of the same, and the crawfish discharges it by way of the gills; and, by the way, the gill-shaped organs in the crawfish are very numerous.
The following properties are common to all crustaceans: they have in all cases two teeth, or mandibles (for the front teeth in the crawfish are two in number), and in all cases there is in the mouth a small fleshy structure serving for a tongue; and the stomach is close to the mouth, only that the crawfish has a little oesophagus in front of the stomach, and there is a straight gut attached to it. This gut, in the crawfish and its congeners, and in the carids, extends in a straight line to the tail, and terminates where the animal discharges the residuum, and where the female deposits her spawn; in the crab it terminates where the flap is situated, and in the centre of the flap. (And by the way, in all these animals the spawn is deposited outside.) Further, the female has the place for the spawn running along the gut. And, again, all these animals have, more or less, an organ termed the ‘mytis’, or ‘poppyjuice’.
We must now proceed to review their several differentiae.
The crawfish then, as has been said, has two teeth, large and hollow, in which is contained a juice resembling the mytis, and in between the teeth is a fleshy substance, shaped like a tongue. After the mouth comes a short oesophagus, and then a membranous stomach attached to the oesophagus, and at the orifice Of the stomach are three teeth, two facing one another and a third standing by itself underneath. Coming off at a bend from the stomach is a gut, simple and of equal thickness throughout the entire length of the body until it reaches the anal vent.
These are all common properties of the crawfish, the carid, and the crab; for the crab, be it remembered, has two teeth.
Again, the crawfish has a duct attached all the way from the chest to the anal vent; and this duct is connected with the ovary in the female, and with the seminal ducts in the male. This passage is attached to the concave surface of the flesh in such a way that the flesh is in betwixt the duct and the gut; for the gut is related to the convexity and this duct to 1087
the concavity, pretty much as is observed in quadrupeds. And the duct is identical in both the sexes; that is to say, the duct in both is thin and white, and charged with a sallow-coloured moisture, and is attached to the chest.
(The following are the properties of the egg and of the convolutes in the carid.)
The male, by the way, differs from the female in regard to its flesh, in having in connexion with the chest two separate and distinct white substances, resembling in colour and conformation the tentacles of the cuttle-fish, and they are convoluted like the ‘poppy’ or quasi-liver of the trumpet-shell. These organs have their starting-point in ‘cotyledons’ or papillae, which are situated under the hindmost feet; and hereabouts the flesh is red and blood-coloured, but is slippery to the touch and in so far unlike flesh. Off from the convolute organ at the chest branches off another coil about as thick as ordinary twine; and underneath there are two granular seminal bodies in juxta-position with the gut. These are the organs of the male. The female has red-coloured eggs, which are adjacent to the stomach and to each side of the gut all along to the fleshy parts, being enveloped in a thin membrane.
Such are the parts, internal and external, of the carid.
<
div id="section59" class="section" title="3"> 3
The inner organs of sanguineous animals happen to have specific designations; for these animals have in all cases the inner viscera, but this is not the case with the bloodless animals, but what they have in common with red-blooded animals is the stomach, the oesophagus, and the gut.
With regard to the crab, it has already been stated that it has claws and feet, and their position has been set forth; furthermore, for the most part they have the right claw bigger and stronger than the left. It has also been stated’ that in general the eyes of the crab look sideways. Further, the trunk of the crab’s body is single and undivided, including its head and any other part it may possess. Some crabs have eyes placed sideways on the upper part, immediately under the back, and standing a long way apart, and some have their eyes in the centre and close together, like the crabs of Heracleotis and the so-called ‘grannies’. The mouth lies underneath the eyes, and inside it there are two teeth, as is the case with the crawfish, only that in the crab the teeth are not rounded but long; and over the teeth are two lids, and in betwixt them are structures 1088
such as the crawfish has besides its teeth. The crab takes in water near by the mouth, using the lids as a check to the inflow, and discharges the water by two passages above the mouth, closing by means of the lids the way by which it entered; and the two passage-ways are underneath the eyes. When it has taken in water it closes its mouth by means of both lids, and ejects the water in the way above described. Next after the teeth comes the oesophagus, very short, so short in fact that the stomach seems to come straightway after the mouth. Next after the oesophagus comes the stomach, two-horned, to the centre of which is attached a simple and delicate gut; and the gut terminates outwards, at the operculum, as has been previously stated. (The crab has the parts in between the lids in the neighbourhood of the teeth similar to the same parts in the crawfish.) Inside the trunk is a sallow juice and some few little bodies, long and white, and others spotted red. The male differs from the female in size and breadth, and in respect of the ventral flap; for this is larger in the female than in the male, and stands out further from the trunk, and is more hairy (as is the case also with the female in the crawfish).
So much, then, for the organs of the malacostraca or crustacea.
<
div id="section60" class="section" title="4"> 4
With the ostracoderma, or testaceans, such as the land-snails and the sea-snails, and all the ‘oysters’ so-called, and also with the sea-urchin genus, the fleshy part, in such as have flesh, is similarly situated to the fleshy part in the crustaceans; in other words, it is inside the animal, and the shell is outside, and there is no hard substance in the interior. As compared with one another the testaceans present many diversities both in regard to their shells and to the flesh within. Some of them have no flesh at all, as the sea-urchin; others have flesh, but it is inside and wholly hidden, except the head, as in the land-snails, and the so-called cocalia, and, among pelagic animals, in the purple murex, the ceryx or trumpet-shell, the sea-snail, and the spiral-shaped testaceans in general.
Of the rest, some are bivalved and some univalved; and by ‘bivalves’ I mean such as are enclosed within two shells, and by ‘univalved’ such as are enclosed within a single shell, and in these last the fleshy part is exposed, as in the case of the limpet. Of the bivalves, some can open out, like the scallop and the mussel; for all such shells are grown together on one side and are separate on the other, so as to open and shut. Other bivalves are closed on both sides alike, like the solen or razor-fish. Some 1089
testaceans there are, that are entirely enveloped in shell and expose no portion of their flesh outside, as the tethya or ascidians.
Again, in regard to the shells themselves, the testaceans present differences when compared with one another. Some are smooth-shelled, like the solen, the mussel, and some clams, viz. those that are nicknamed
‘milkshells’, while others are rough-shelled, such as the pool-oyster or edible oyster, the pinna, and certain species of cockles, and the trumpet shells; and of these some are ribbed, such as the scallop and a certain kind of clam or cockle, and some are devoid of ribs, as the pinna and another species of clam. Testaceans also differ from one another in regard to the thickness or thinness of their shell, both as regards the shell in its entirety and as regards specific parts of the shell, for instance, the lips; for some have thin-lipped shells, like the mussel, and others have thick-lipped shells, like the oyster. A property common to the above mentioned, and, in fact, to all testaceans, is the smoothness of their shells inside. Some also are capable of motion, like the scallop, and indeed some aver that scallops can actually fly, owing to the circumstance that they often jump right out of the apparatus by means of which they are caught; others are incapable of motion and are attached fast to some external object, as is the case with the pinna. All the spiral-shaped testaceans can move and creep, and even the limpet relaxes its hold to go in quest of food. In the case of the univalves and the bivalves, the fleshy substance adheres to the shell so tenaciously that it can only be removed by an effort; in the case of the stromboids, it is more loosely attached. And a peculiarity of all the stromboids is the spiral twist of the shell in the part farthest away from the head; they are also furnished from birth with an operculum. And, further, all stromboid testaceans have their shells on the right hand side, and move not in the direction of the spire, but the opposite way. Such are the diversities observed in the external parts of these animals.
The internal structure is almost the same in all these creatures, and in the stromboids especially; for it is in size that these latter differ from one another, and in accidents of the nature of excess or defect. And there is not much difference between most of the univalves and bivalves; but, while those that open and shut differ from one another but slightly, they differ considerably from such as are incapable of motion. And this will be illustrated more satisfactorily hereafter.
The spiral-shaped testaceans are all similarly constructed, but differ from one another, as has been said, in the way of excess or defect (for the larger species have larger and more conspicuous organs, and the smaller 1090
have smaller and less conspicuous), and, furthermore, in relative hardness or softness, and in other such accidents or properties. All the stromboids, for instance, have the flesh that extrudes from the mouth of the shell, hard and stiff; some more, and some less. From the middle of this protrudes the head and two horns, and these horns are large in the large species, but exceedingly minute in the smaller ones. The head protrudes from them all in the same way; and, if the animal be alarmed, the head draws in again. Some of these creatures have a mouth and teeth, as the snail; teeth sharp, and small, and delicate. They have also a proboscis just like that of the fly; and the proboscis is tongue-shaped. The ceryx and the purple murex have this organ firm and solid; and just as the myops, or horse-fly, and the oestrus, or gadfly, can pierce the skin of a quadruped, so is that proboscis proportionately stronger in these testaceans; for they bore right through the shells of other shell-fish on which they prey. The stomach follows close upon the mouth, and, by the way, this organ in the snail resembles a bird’s crop. Underneath come two white firm formations, mastoid or papillary in form; and similar formations are found in the cuttle-fish also, only that they are of a firmer consistency in the cuttle-fish. After the stomach comes an oesophagus, simple and long, extending to the poppy or quasi-liver, which is in the innermost recess of the shell. All these statements may be verified in the case of the purple murex and the ceryx by observation within the whorl of the shell. What comes next to the oesophagus is the gut; in fact, the gut is continuous with the oesophagus, and runs its whole length uncomplicated to the outlet of the residuum. The gut has its point of origin in the region of the coil of the mecon, or so-called ‘poppy’, and is wider hereabouts (for remember, the mecon is for the most part a sort of excretion in all testaceans); it then takes a bend and runs up again towards the fleshy part, and terminates by the side of the head, where the animal discharges its residuum; and this holds good in the case of all stromboid testaceans, whether terrestrial or marine. From the stomach there is drawn in a parallel direction with the oesophagus, in the larger snails, a long white duct enveloped in a membrane, resembling in colour the mastoid formations higher up; and in it are nicks or interruptions, as in the egg-mass of the crawfish, only, by the way, the duct of which we are treating is white and the egg-mass of the crawfish is red. This formation has no outlet nor duct, but is enveloped in a thin membrane with a narrow cavity in its interior. And from the gut downward extend black and rough formations, in close connexion, something like the formations in the tortoise, only not so black. Marine snails, also, have these formations, 1091
and the white ones, only that the formations are smaller in the smaller species.
The non-spiral univalves and bivalves are in some respect similar in construction, and in some respects dissimilar, to the spiral testaceans.
They all have a head and horns, and a mouth, and the organ resembling a tongue; but these organs, in the smaller species, are indiscernible owing to the minuteness of these animals, and some are indiscernible even in the larger species when dead, or when at rest and motionless. They all have the mecon, or poppy, but not all in the same place, nor of equal size, nor similarly open to observation; thus, the limpets have this organ deep down in the bottom of the shell, and the bivalves at the hinge connecting the two valves. They also have in all cases the hairy growths or beards, in a circular form, as in the scallops. And, with regard to the so-called ‘egg’, in those that have it, when they have it, it is situated in one of the semi-circles of the periphery, as is the case with the white formation in the snail; for this white formation in the snail corresponds to the so-called egg of which we are speaking. But all these organs, as has been stated, are distinctly traceable in the larger species, while in the small ones they are in some cases almost, and in others altogether, indiscernible. Hence they are most plainly visible in the large scallops; and these are the bivalves that have one valve flat-shaped, like the lid of a pot. The outlet of the excretion is in all these animals (save for the exception to be afterwards related) on one side; for there is a passage whereby the excretion passes out. (And, remember, the mecon or poppy, as has been stated, is an excretion in all these animals-an excretion enveloped in a membrane.) The so-called egg has no outlet in any of these creatures, but is merely an excrescence in the fleshy mass; and it is not situated in the same region with the gut, but the ‘egg’ is situated on the right-hand side and the gut on the left. Such are the relations of the anal vent in most of these animals; but in the case of the wild limpet (called by some the ‘sea-ear’), the residuum issues beneath the shell, for the shell is perforated to give an outlet. In this particular limpet the stomach is seen coming after the mouth, and the egg-shaped formations are discernible. But for the relative positions of these parts you are referred to my Treatise on Anatomy.
The so-called carcinium or hermit crab is in a way intermediate between the crustaceans and the testaceans. In its nature it resembles the crawfish kind, and it is born simple of itself, but by its habit of introducing itself into a shell and living there it resembles the testaceans, and so appears to partake of the characters of both kinds. In shape, to give a 1092
simple illustration, it resembles a spider, only that the part below the head and thorax is larger in this creature than in the spider. It has two thin red horns, and underneath these horns two long eyes, not retreating inwards, nor turning sideways like the eyes of the crab, but protruding straight out; and underneath these eyes the mouth, and round about the mouth several hair-like growths, and next after these two bifurcate legs or claws, whereby it draws in objects towards itself, and two other legs on either side, and a third small one. All below the thorax is soft, and when opened in dissection is found to be sallow-coloured within. From the mouth there runs a single passage right on to the stomach, but the passage for the excretions is not discernible. The legs and the thorax are hard, but not so hard as the legs and the thorax of the crab. It does not adhere to its shell like the purple murex and the ceryx, but can easily slip out of it. It is longer when found in the shell of the stromboids than when found in the shell of the neritae.
And, by the way, the animal found in the shell of the neritae is a separate species, like to the other in most respects; but of its bifurcate feet or claws, the right-hand one is small and the left-hand one is large, and it progresses chiefly by the aid of this latter and larger one. (In the shells of these animals, and in certain others, there is found a parasite whose mode of attachment is similar. The particular one which we have just described is named the cyllarus.)
The nerites has a smooth large round shell, and resembles the ceryx in shape, only the poppy-juice is, in its case, not black but red. It clings with great force near the middle. In calm weather, then, they go free afield, but when the wind blows the carcinia take shelter against the rocks: the neritae themselves cling fast like limpets; and the same is the case with the haemorrhoid or aporrhaid and all others of the like kind. And, by the way, they cling to the rock, when they turn back their operculum, for this operculum seems like a lid; in fact this structure represents the one part, in the stromboids, of that which in the bivalves is a duplicate shell. The interior of the animal is fleshy, and the mouth is inside. And it is the same with the haemorrhoid, the purple murex, and all suchlike animals.
Such of the little crabs as have the left foot or claw the bigger of the two are found in the neritae, but not in the stromboids. are some snail-shells which have inside them creatures resembling those little crayfish that are also found in fresh water. These creatures, however, differ in having the part inside the shells But as to the characters, you are referred to my Treatise on Anatomy.
<
1093
div id="section61" class="section" title="5"> 5
The urchins are devoid of flesh, and this is a character peculiar to them; and while they are in all cases empty and devoid of any flesh within, they are in all cases furnished with the black formations. There are several species of the urchin, and one of these is that which is made use of for food; this is the kind in which are found the so-called eggs, large and edible, in the larger and smaller specimens alike; for even when as yet very small they are provided with them. There are two other species, the spatangus, and the so-called bryssus, these animals are pelagic and scarce. Further, there are the echinometrae, or ‘mother-urchins’, the largest in size of all the species. In addition to these there is another species, small in size, but furnished with large hard spines; it lives in the sea at a depth of several fathoms; and is used by some people as a specific for cases of strangury. In the neighbourhood of Torone there are sea-urchins of a white colour, shells, spines, eggs and all, and that are longer than the ordinary sea-urchin. The spine in this species is not large nor strong, but rather limp; and the black format