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The Brain:

Understanding Neurobiology

Through the Study of Addiction

under a contract from the

National Institutes of Health

U.S. Department of Health and Human Services

National Institutes of Health

National Institute on Drug Abuse

Center for Curriculum Development

5415 Mark Dabling Boulevard

Colorado Springs, CO 80918

BSCS Development Team

William Mowczko, Office of Science Education

Nancy M. Landes, Principal Investigator

Cathrine Sasek, National Institute on Drug Abuse

Anne L. Westbrook, Project Director

Gloria Seelman, Office of Science Education

Debra A. Hannigan, Curriculum Developer

Field-test Teachers

Ann C. Lanari, Research Assistant

Domenica Altieri, Sonora High School, La Habra, California

Carol Vallee, Project Assistant

William Barlow, Preston High School, Kingwood, West Virginia

Karen Bertollini, Project Assistant

Gwyn Bush, McNicholas High School, Cincinnati, Ohio

Mary Crist, Project Assistant

Jennifer Carpio, St. Joseph’s Academy, St. Louis, Missouri

Carrie Hamm, Project Assistant

Kathy Cattrell, Crestview High School, Columbiana, Ohio

Raphaela Conner, Project Assistant

Aster Chin, Lowell High School, San Francisco, California

Barbara Perrin, Production Manager

Katy Colvin, Fort LeBoeuf High School, Waterford, Pennsylvania

Ric Bascobert, Editor

Linda Dizer, Girls Preparatory School, Chattanooga, Tennessee

Diane Gionfriddo, Photo Research

Karen Emery, Shady Spring High School, Shady Spring, West Virginia

Lisa Chilberg, Graphic Designer

Fran Enright, Evergreen Senior High School, Evergreen, Colorado

Sandra Matthews, Evaluator

Leon Fox, Perry High School, Perry, Iowa

Videodiscovery, Inc., Development Team

Charlotte Freeman, Girls Preparatory School, Chattanooga, Tennessee

Shaun Taylor, Vice President for Product Development

Marian Gonzalez, Lowell High School, San Francisco, California

Michael Bade, Multimedia Producer/Project Director/Videographer

Paula Henderson, Newark High School, Wilmington, Delaware

Greg Humes, Assistant Multimedia Producer

Wade Hill, Battle Mountain High School, Minturn, Colorado

Cathy Saum, Graphic Designer

John Johnson, Creekview High School, Carrollton, Texas

Lucy Flynn Zuccotti, Photo Research

Karel Lilly, Foshay Learning Center, Los Angeles, California

Michele Moore, Assistant to Project Director

Marilyn Link, North Carolina School of Science and Mathematics,

Jennifer Hunt, Costumes and Makeup

Durham, North Carolina

Mike Commins, Gaffer (Lighting)

Brian McCarry, Monsignor Bonner High School, Drexel Hill,

Chet McKnight, Sound

Pennsylvania

Lee Strucker, Script Writer

Glenn Miller, Wheaton High School, Wheaton, Maryland

Tracy Peterson, Hamilton East High School, Hamilton, New Jersey

Advisory Committee

Kathleen Ranwez, Pomona High School, Arvada, Colorado

Andrea Baruchin, Vanderbilt University, Nashville, Tennessee

Ruth Regent-Smith, Pius XI High School, Milwaukee, Wisconsin

Michael Dougherty, Hampden-Sydney College,

Martin Shields, James Caldwell High School, West Caldwell,

Hampden-Sydney, Virginia

New Jersey

David Friedman, Wake Forest University, Winston-Salem,

Ann Sowd, Jackson High School, Massillon, Ohio

North Carolina

Sandra Sundlof, Wheaton High School, Wheaton, Maryland

Reese Jones, University of California, San Francisco, California

Kay Thornton, Perry High School, Perry, Iowa

Kathleen Ranwez, Pomona High School, Arvada, Colorado

Linda Wright, Creekview High School, Carrollton, Texas

Cathrine Sasek, National Institute on Drug Abuse, Bethesda, Maryland

Lois Wysocki, Radnor High School, Radnor, Pennsylvania

Martin Shields, James Caldwell High School, West Caldwell, New Jersey

Susan Wooley, American School Health Association, Kent, Ohio

Special Thanks

BSCS and Videodiscovery thank the following individuals for

Writing Team

providing resources or reviewing specific activities in this unit:

Mary Ann Cutter, University of Colorado, Colorado Springs, Colorado

Dr. William Armstrong, University of Tennessee;

JoAnne Dombrowski, Consultant, Somerton, Arizona

Dr. Richard Cannon, National Heart, Lung and Blood Institute;

Michael Dougherty, Hampden-Sydney College,

Dr. Monica Skarulis, National Institute of Diabetes and Digestive

Hampden-Sydney, Virginia

and Kidney Diseases;

David Friedman, Wake Forest University, Winston-Salem,

Dr. Michael Phelps, UCLA School of Medicine;

North Carolina

Dr. Johannes Czernin, UCLA School of Medicine;

Paula Henderson, Newark High School, Wilmington, Delaware

David Twomey, UCLA School of Medicine; and

Laura McNicholas, Veterans Affairs Medical Center,

Jim Strommer, UCLA School of Medicine.

Philadelphia, Pennsylvania

E. Leong Way, University of California San Francisco,

Cover Image Description

San Francisco, California

The cover shows a positron emission tomography (PET) image of

a human brain. Blood flow to a particular brain area, the amygdala,

Artist

increases when a person addicted to cocaine experiences cravings

Susan Bartel

for the drug. The image, when compared with those taken of people

who aren’t addicted to cocaine, reveals that just eliciting memories

Cover Design

of drug abuse in the addicted person is sufficient to cause changes

Martha Blalock, Medical Arts and Photography Branch,

in brain activity.

National Institutes of Health

This material is based on work supported by the National Institutes

Cover Illustration

of Health under Contract No. 263-98-C-0056. Any opinions, find-

Anna Rose Childress, Ph.D., University of Pennsylvania

ings, conclusions, or recommendations expressed in this publication

School of Medicine

are those of the authors and do not necessarily reflect the view of

Design and Layout

the funding agency.

Angela Greenwalt, Finer Points Productions

Fourth printing, 2010. Copyright © 2000 by BSCS and Videodis-

Photo Credits

covery, Inc. Third printing, 2004; second printing, 2003. All rights

Carlye Calvin

reserved. You have the permission of BSCS and Videodiscovery,

Inc., to reproduce items in this module for your classroom use.

BSCS Administrative Staff

The copyright on this module, however, does not cover reproduc-

Carlo Parravano, Chairman, Board of Directors

tion of these items for any other use. For permissions and other

Rodger W. Bybee, Executive Director

rights under this copyright, please contact BSCS, 5415 Mark

Janet Carlson Powell, Associate Director, Chief Science

Dabling Blvd., Colorado Springs, CO 80918-3842; www.bscs.org;

Education Officer

info@bscs.org; (719) 531-5550.

Larry Satkowiak, Associate Director, Chief Operating Officer

Revised and reprinted March 2010

Videodiscovery, Inc., Administrative Staff

NIH Publication No. 09-4871 ISBN: 1-929614-05-5

D. Joseph Clark, President

Shaun Taylor, Vice President for Product Development

Please contact the NIH Office of Science

National Institutes of Health

Education with questions about this supplement

Bruce Fuchs, Office of Science Education

Lucinda Miner, National Institute on Drug Abuse

at supplements@science.education.nih.gov.

Contents

Foreword. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . v About the National Institutes of Health . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . vii About the National Institute on Drug Abuse . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ix The Essence of Drug Addiction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . xi Introduction to the Module . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1

What Are the Objectives of the Module? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .1

Why Teach the Module? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .1

What’s in It for the Teacher? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2

Implementing the Module . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

What Are the Goals of the Module? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .3

What Are the Science Concepts and How Are They Connected? . . . . . . . . . . . . . . . . . . . . . . . . . . . . .3

How Does the Module Correlate with the National Science Education Standards?. . . . . . . . . . . . . . . . .4

Teaching Standards. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

Assessment Standards . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

How Does the BSCS 5E Instructional Model Promote Active, Collaborative,

Inquiry-Based Learning? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .4

Engage . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6

Explore . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6

Explain . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6

Elaborate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7

Evaluate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7

How Does the Module Support Ongoing Assessment? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .10

How Can Controversial Topics Be Handled in the Classroom? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .10

Using the Web Site . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13

Hardware and Software Requirements. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .13

Getting the Most Out of the Web Site . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .13

Collaborative Groups . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .14

Web Activities for People with Disabilities . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .14

Using the Student Lessons . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15

Format of the Lessons . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .15

Timelines for Teaching the Module . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .16

Student Lessons

Lesson 1— The Brain: What’s Going On in There? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .19

Lesson 2— Neurons, Brain Chemistry, and Neurotransmission . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .41

Lesson 3— Drugs Change the Way Neurons Communicate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .65

Lesson 4— Drug Abuse and Addiction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91

Lesson 5— Drug Addiction Is a Disease, So What Do We Do about It? . . . . . . . . . . . . . . . . . . . . . . . . .125

iii

Additional Resources for Teachers . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143

Glossary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145

References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 153

Masters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 157

iv

Foreword

This curriculum supplement, from The NIH

Activities promote active and collaborative learning

Curriculum Supplement Series, brings cutting-edge

and are inquiry-based to help students develop

medical science and basic research discoveries

problem-solving strategies and critical thinking.

from the laboratories of the National Institutes

of Health (NIH) into classrooms. As the largest

Each curriculum supplement comes with a complete

medical research institution in the United

set of materials for both teachers and students,

States, NIH plays a vital role in the health of

including printed materials, extensive background

all Americans and seeks to foster interest in

and resource information, and a Web site with videos

research, science, and medicine-related careers

and interactive activities. The supplements are

for future generations. NIH’s Office of Science

distributed at no cost to teachers across the United

Education (OSE) is dedicated to promoting

States. All materials may be copied for classroom use

science education and scientific literacy.

but may not be sold.

We designed this curriculum supplement to

For a complete list of curriculum supplements,

complement existing life science curricula at both

updates, availability, and ordering information, or

the state and local levels and to be consistent

to submit feedback, please visit our Web site or

with the National Science Education Standards.1 It

write to

was developed and tested by a team composed of

Curriculum Supplement Series

teachers, scientists, medical experts, and other

Office of Science Education

professionals with relevant subject-area expertise

National Institutes of Health

from institutes and medical schools across

6100 Executive Boulevard

the country, representatives from the National

Suite 3E01 700 MSC 7520

Institute on Drug Abuse, and curriculum design

Bethesda, MD 20892-7520

experts from Biological Sciences Curriculum

Study (BSCS) and Videodiscovery. The authors

We appreciate the valuable contributions of the

incorporated real scientific data and actual case

talented staff at Biological Sciences Curriculum

studies into classroom activities. A three-year

Study (BSCS) and Videodiscovery, Inc. We are also

development process included geographically

grateful to the NIH scientists, advisors, and all other

dispersed field tests by teachers and students.

participating professionals for their work and

For the 2010 (fourth) printing, key sections of

dedication. Finally, we thank the teachers and

the supplement were updated, but the Student

students who participated in focus groups and field

Lessons remain basically the same.

tests to ensure that these supplements are both

engaging and effective. I hope you find our series a

The structure of this module enables teachers

valuable addition to your classroom and wish you a

to effectively facilitate learning and stimulate

productive school year. We welcome your feedback.

student interest by applying scientific concepts

to real-life scenarios. Design elements include

Bruce A. Fuchs, Ph.D.

a conceptual flow of lessons based on the BSCS

Director

5E Instructional Model (page 3), multisubject

Office of Science Education

integration emphasizing cutting-edge science

National Institutes of Health

content, and built-in assessment tools.

supplements@science.education.nih.gov

________________________

1 The National Academy of Sciences released the National Science Education Standards in 1996, outlining what all citizens should understand about science by the time they graduate from high school. The Standards encourages teachers to select major science concepts that empower students to use information to solve problems rather than stressing memorization of unrelated information.

v

About the National Institutes of Health

Begun as the one-room Laboratory of Hygiene

libraries and the training of medical librarians

in 1887, the National Institutes of Health (NIH)

and other health information specialists.

today is one of the world’s foremost biomedical

and behavioral research centers and the federal

Organization

focal point for health research in the United States.

Composed of 27 separate institutes and centers, NIH

is one of eight health agencies of the Public Health

Mission and Goals

Service within the U.S. Department of Health and

The NIH mission is science in pursuit of

Human Services. NIH encompasses 75 buildings

fundamental knowledge about the nature and

on more than 300 acres in Bethesda, Md., as well

behavior of living systems and the application

as facilities at several other sites in the United

of that knowledge to extend healthy life and

States. The NIH budget has grown from about

reduce the burdens of illness and disability.

$300 in 1887 to more than $30 billion in 2009.

The goals of the agency are to

• foster fundamental creative discoveries

Research Programs

and innovative research strategies and

One of NIH’s principal concerns is to invest wisely

their applications as a basis for advancing

the tax dollars entrusted to it for the support and

significantly the nation’s capacity to protect

conduct of this research. Approximately 82 percent

and improve health;

of the investment is made through grants and

• develop, maintain, and renew scientific

contracts supporting research and training in more

resources—both human and physical—that will

than 2,000 research institutions throughout the

ensure the nation’s ability to prevent disease;

United States and abroad. In fact, NIH grantees

• expand the knowledge base in medical and

are located in every state in the country. These

associated sciences in order to enhance the

grants and contracts make up the NIH Extramural

nation’s economic well-being and ensure a

Research Program.

continued high return on the public investment

in research; and

Approximately 10 percent of the budget goes to

• exemplify and promote the highest level of

NIH’s Intramural Research Programs, the more

scientific integrity, public accountability, and

than 2,000 projects conducted mainly in its own

social responsibility in the conduct of science.

laboratories. These projects are central to the NIH

scientific effort. First-rate intramural scientists

NIH works toward meeting those goals by providing

collaborate with one another regardless of institute

leadership, direction, and grant support to programs

affiliation or scientific discipline and have the

designed to improve the health of the nation

intellectual freedom to pursue their research leads

through research into the

in NIH’s own laboratories. These explorations

• causes, diagnosis, prevention, and cure of

range from basic biology to behavioral research,

human diseases;

to studies of treatments for major diseases.

• processes of human growth and development;

• biological effects of environmental

Grant-Making Process

contaminants;

The grant-making process begins with an idea

• understanding of mental, addictive, and

that an individual scientist describes in a written

physical disorders; and

application for a research grant. The project

• collection, dissemination, and exchange of

might be small, or it might involve millions

information in medicine and health, including

of dollars. The project might become useful

the development and support of medical

immediately as a diagnostic test or new treatment,

vii

or it might involve studies of basic biological or

• In 1990, NIH researchers performed the first

behavioral processes whose clinical value may not

trial of gene therapy in humans. Scientists

be apparent for many years.

are increasingly able to locate, identify, and

describe the functions of many of the genes

Each research grant application undergoes peer

in the human genome. The ultimate goal is to

review. A panel of scientific experts, primarily

develop screening tools and gene therapies for

from outside the government, who are active and

the general population for cancer and many

productive researchers in the health sciences first

other diseases.

evaluates the scientific merit of the application.

Then, a national advisory council or board,

Science Education

composed of eminent scientists as well as

Science education by NIH and its institutes and

members of the public who are interested in

centers contributes to ensuring the continued

health issues or the biomedical or behavioral

supply of well-trained basic research and clinical

sciences, determines the project’s overall merit

investigators, as well as the myriad professionals

and priority in advancing the research agenda of

in the many allied disciplines who support the

the particular NIH funding institutes and centers.

research enterprise. These efforts also help

educate people about scientific results so that

About 38,500 research and training applications

they can make informed decisions about their

are reviewed annually throughout the NIH peer-

own—and the public’s—health.

review system. At any given time, NIH supports

35,000 grants in universities, medical schools, and

This curriculum supplement is one such science

other research and research training institutions,

education effort, a collaboration among four

both nationally and internationally.

partners: the NIH National Institute on Drug

Abuse, the NIH Office of Science Education